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Steven G. Boxer

Camille and Henry Dreyfus Professor of Chemistry, Biophysical and Physical Chemistry, Chemistry, Stanford University


Telephone: (650)723-4482
Fax: (650)723-4817
Email: | Web site

Stanford University, Stanford, CA (b. 1947). B.S. with honors 1969, Tufts University; Ph.D. 1976, University of Chicago. Phone: (650)723-4482, Fax: (650)723-4817, email:

Boxer works on structure and function in several biological systems with applications in biotechnology. Research topics include the mechanism of the initial events in photosynthesis, electrostatics and dynamics in heme proteins, physical chemistry of supported lipid bilayers, including the construction of interfaces between biological membranes, cells, and non-biological surfaces, and development of Stark spectroscopy, which is applicable to a wide range of molecules and types of transitions. Approaches include ultrafast-to-ultraslow absorption and fluorescence, magnetic resonance, FTIR, microscopy, photolithography, and recombinant DNA methods.

Selected Publications:

  1. Yoshina-Ishii, C., Miller, G. P., Kraft, M. L. Kool, E. T., Boxer, S. G.; "General Method for Modification of Liposomes for Encoded Assembly on Supported Bilayers". Journal of the American Chemical Society, 127, 1356-1357, 2005.
  2. Kraft, M. L., Weber, P. K., Longo, M. L., Boxer, S. G., Hutcheon, I. D., "Phase Separation of Lipid Membranes Analyzed with High-Resolution Secondary-Ion Mass Spectrometry", Science, 313, 1948-1951, 2006.
  3. Tanaka, M., Hermann, J., Haase, I., Fischer, M., Boxer, S.; "Frictional Drag and Electrical Manipulation of Recombinant Proteins in Polymer Supported Membranes". Langmuir, 23, 5638-5644, 2007.


    Chan, Y.-H., Lenz, P., Boxer, S.; "Kinetics of DNA-Mediated Docking Reactions between Vesicles Tethered to Supported Lipid Bilayers". Proceedings of the National Academy of Sciences, 104, 18913-18918, 2007.


    ChanY.-H., Boxer,S. "Model Membrane Systems and their Applications". Current Opinion in Chemical Biology, 11, 581-587, 2007.


    Lin, W-C., Blanchette, C. D., Longo, M.L.; “Fluid-Phase Chain Unsaturation Controlling Domain Microstructure and Phase in Ternary Lipid Bilayers Containing GalCer and Cholesterol”. Biophysical Journal, 92, 2831-2841, 2007.


    Blanchette, C. D., Lin, W-C., Orme, C. A., Ratto, T. V., Longo, M. L., “Using Nucleation Rates to Determine Interfacial Line Tension of Symmetric and Asymmetric Lipid Bilayer Domains”. Langmuir, 23, 5875-5877, 2007.


    Blanchette, C. D., Orme, C. A., Ratto, T. V., Longo, M. L.; “Quantifying Growth of Symmetric and Asymmetric Lipid Bilayer Domains”. Molecular and Surface Forces, Langmuir Special Issue, 24, 1219-1224, 2008.


    Blanchette, C. D., Lin, W-C., Orme, C. A., Ratto, T. V, Longo, M. L., “Domain Nucleation Rates and Interfacial Line Tensions in Supported Bilayers of Ternary Mixtures Containing Galactosylceramide”. Biophysical Journal, 94, 2691–2697, 2008.


    Kasson, P., Pande, V. S.; “Predicting Structure and Dynamics of Loosely-Ordered Protein Complexes: Influenza Hemagglutinin Fusion Peptide.” Pacific Symposium on Biocomputing, 12, 40-50, 2007.


    Teclemariam, N.P., Beck, V.A., Shaqfeh, E.S.G., Muller, S.J.; “Dynamics of DNA Polymers in Post Arrays: Comparison of Single Molecule Experiments and Simulations”. Macromolecules, 40, 3848-3859, 2007.

    Lueth, C.A., Shaqfeh, E.S.G., “Tethered DNA Shear Dynamics in the Flow Gradient Plane: Application to Double Tethering”. Korea-Australia J. Rheol., 19, 3, 141-146, 2007.


    Beck, V.A., Shaqfeh, E.S.G., “Ergodicity-Breaking and the Unravelling Dynamics of a Polymer in Linear and Non-Linear Extensional Flows”. J. Rheol., 51, 3, 561-574, May/June, 2007.


    Hoffman, B., Shaqfeh, E.S.G., “The Dynamics of the Coil-Stretch Transition for Long, Flexible Polymers in Planar Mixed Flows''. J. Rheol., 51, 5, 947-969, 2007 (see also Virtual Journal of Biological Physics Research September 1, 2007.)